DNA replication

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Slide 1 : APT ACADEMIC SOLUTIONS believes in people who will deliver the right solutions, and it proudly possesses them.

Home Work : Home Work Give examples in which single stranded DNA & double stranded RNA occurs Single stranded DNA occurs in f174 Double stranded RNA occurs in Cauliflower virus

Slide 5 : Session Objectives DNA Replication Transcription

Slide 6 : Replication of DNA Self duplication process of DNA Semi-conservative in nature Replication produce molecules with both old and new DNA Each molecule consists of one old strand and one new one, hence called as semi-conservative

Slide 7 : Experimental evidence for semi-conservative replication: Meselson and Stahl (1958) provided experimental evidence supporting semi-conservative mode of DNA replication in E. coli by DNA banding techinique Taylor provided experimental evidence supporting semi-conservative mode of DNA replication in Vicia faba by autoradiographic technique Semi-conservative replication of DNA

DNA Replication : DNA Replication Replication occurs in replication fork or replication bubble Prokaryotes are mono repliconic Eukaryotes are multi-repliconic

Step 1: Unwinding of DNA chain : Step 1: Unwinding of DNA chain

Illustrative Problem : Illustrative Problem Unwinding of DNA is done by topoisomerases exonuclease helicase ligase DNA is helical structure & for unwinding negative coiling has to be induced. Topoisomerase is known for inducing negative coiling or unwinding of DNA chain.

Step 2: Priming : Step 2: Priming

Step 3: Polymerization : Step 3: Polymerization DNA polymerase is the main replicating enzyme Complimentary base pairing occurs Chain grows in 5’ to 3’ direction

Slide 13 : Step 4: Leading & Lagging strand As a result of further opening of chain, one strand has continuous synthesis and is known as leading strand Discontinuous synthesis on other strand & is known as lagging strand Okazaki fragments are formed on lagging strand Lagging strand is formed as synthess always occurs in 5’ to 3’ direction

Step 5: DNA Polymerase I : Step 5: DNA Polymerase I It act as proof reader Removes the wrongly inserted base pair It fills the gap formed due to removal of base pair by complimentary base pairing

Slide 15 : Step 6: DNA Ligase DNA Ligase seals the gap It forms PDE bond It is NAD dependent enzyme

Step 7: Topoisomerase (DNA Gyrase) : Step 7: Topoisomerase (DNA Gyrase)

DNA Replication : Prokaryote vs Eukaryote : DNA Replication : Prokaryote vs Eukaryote

Quiz : Quiz Leading & lagging strands are formed simultaneously in both strand of a replicon True / False

Central Dogma : Central Dogma Coined by Crick (1958) It is flow of information from genetic material to protein synthesis Involves two steps – Transcription & Translation DNA Transcription m-RNA Translation Protein

Reverse Transcription (Temins & Baltimore) : Reverse Transcription (Temins & Baltimore) Genetic material of retrovirus (like HIV)is RNA It uses its genetic material for the formation of DNA Process is known as reverse transcription Enzyme used in the mechanism is reverse transcriptase m-RNA Reverse Transcription DNA Transcripton m-RNA Protein

Transcription : Transcription Process of formation of RNA from DNA template Involves rewriting the genetic message coded in DNA into an RNA molecule Mechanism can be discussed in 3 steps : Initiation Elongation Termination

Initiation : Initiation Sigma factor disassociate & core enzyme elongates the chain RNA poly binds at P site P site is recognized by sigma factor

Elongation : Elongation DNA chain opens at P site Core enzyme adds complimentary bases Chain grows in 5’ to 3’ direction One chain act as sense strand and other as mis-sence strand

Termination : Termination Core enzyme reaches T site Rho protein is present at T site Entire complex is withdrawn & DNA structure is restored Sigma factor binds with core enz releasing ‘rho’ Rho finally joins T site of DNA

Mutation : Mutation Sudden inheritable changes in genetic material is known as mutation Spontaneous mutation : Occurs randomly & naturally Induced mutation : occurs due to some artificial mutagens

Mutagens : Mutagens

Physical Mutagen : Physical Mutagen Rise in temperature breaks hydrogen bond between the two strands of DNA High energy radiations like X-rays, a, b, g - rays breaks the PDE bond UV rays result in the formation of Thymine dimer

Chemical Mutagen : Chemical Mutagen Nitrous acid is deaminating which changes

Chemical Mutagen : Chemical Mutagen Alkylating agents like Nitrogen mustard, diethyl sulphate, dimethyl nitsoamine etc causes methylation or ethylation of nitrogenous bases. The later fails to pair normally & also prevents separation of DNA strand Base analogous are structurally similar but functionally dissimilar to the normal bases. It disturbs in replication and translation. Acridines enter DNA chain in between two base pairs & cause deletion or addition of few bases. It results in frame shift mutation or gibberish mutation

Mechanism of Mutation : Mechanism of Mutation

Inversion : Inversion Mutagen can change the base sequence of cistron in the reverse order The new sequence will have different codons It will code for different amino acids GCC TAT TTG GTT TAT CCG

Substitution : Substitution Transition : Purine is replaced by another purine (adenine by guanine) Pyrimidine is replaced by another pyrimidine ( cytocine by thymine) Transversion : Purine is replaced or subsituted by a pyrimidine basa and vice versa

Frame-shift Mutation : Frame-shift Mutation Reading of the frame of base sequence shifts laterally either in forward direction due to insertion of base or in backward direction due to deletion of base. Insertion ACC TAT TTG AAC CTA TTT G

Deletion : Deletion Deletion ACC TAT TTG CCT ATT TG One or more segment of DNA is lost

Nonsense Mutation : Nonsense Mutation It stops polypeptide synthesis due to the formation of a terminating codon (UAA, UAG & UGA) Mis-sense Mutation It involves changes in codon resulting into production of non-functional amino acid. Same-sense Mutation

Next Class : Next Class Genetic Code Translation Join me after reading the topics from CBSE book

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